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Tord Snäll

Tord Snäll
We have three main research interests. One core interest is to increase our understanding of, and ability to predict the distribution or dynamics of (meta)populations, species, communities, and ecosystem services. We further investigate how these biodiversity measures and services may respond to scenarios of future land-use or climate change. Finally, we investigate the usefulness of citizen science data for answering basic and applied questions. Most studies concern the use and conservation of forest resources.


Distributions or dynamics of (meta)populations, species or communities

A main research interest is to investigate the relative importance of environmental conditions, species interactions and (meta)population processes, such as dispersal, in explaining the spatial distribution or dynamics of (meta)populations, species and communities. The work on single species includes investigating new approaches for estimating the rate of metapopulation dynamics using very limited amount of data, e.g. only the distribution pattern of a species among its habitat patches. We mainly do statistical modelling of field data on species occurrence or dynamics, but we also conduct field experiments and population genetic studies. Sessile (vascular plants, bryophytes, fungi, lichens) as well as mobile (insects, birds) species are studied, mainly in forests.

Levels and Dynamics of Ecosystem Services

We investigate the relative importance of local environmental conditions, tree species richness and composition in explaining levels and dynamics of ecosystem services on different spatial scales. Provisional, supporting, regulating and cultural services are studied. We answer these questions by statistical modelling of field data, typically large-scale monitoring data, such as national forest inventory data.

Response of species or ecosystem services to scenarios of land-use or climate change

Another main research interest is to investigate how (meta)populations, species, communities and ecosystem services may be affected by scenarios of future forestry or climate change. This includes simulating scenarios (answering What if?) and optimization (answering How?). The main spatial scales studied are the landscape and national scales, but we also consider the EU and global scales, see e.g. Horizon Europe and ERA-Net projects below. The scenarios simulated are formulated together with a range of stakeholders and with our research partners. The questions are answered using models that we develop in the group or available models and tools, e.g. Heureka for forestry planning.

Usefulness of citizen science data for modelling and scenario analysis

The final main research interest is to investigate the usefulness of volunteers’ reports of observations of species to open-access websites, such as Artportalen or GBIF, for answering basic and applied questions. We particularly evaluate them for predicting species distributions, projecting species occurrence into the future, and for population trend estimation.

Funding & researach group

International projects led by us: SustMultBiomass funded by ForestValue.
International projects where we are a partner: ForestNavigator funded by Horizon Europe and ForBioMon funded by Biodiversa+ (starting 2o24).

We also run projects funded by the national organizations Formas and the Swedish Research Council.

Postdocs: Nils Ambec, Laura Chevaux, Federico Lingua, Cristian Montalvo Mancheno.


In our research on land-use, green infrastructure etc. we interact with different stakeholders. See film about this here.

Selected publications

International publications

In press

Probst B, Toraño Caicoya A, Hilmers T, Snäll T, Stoltz J, Suda M. 2024. Predicting the perceived restorativeness and its indicators of present and future forest stands. People and Nature.

Stoltz J, Burgas D, Potterf M, Duflot R, Eyvindson K, Probst BM, Toraño-Caicoya A, Mönkkönen M, Gyllin M, Grahn P, Snäll T. 2024. Forests management for health-promotion: Future developments of salutogenic properties in managed boreal forests. Forests.


Toraño Caicoya A, Vergarechea M, Blattert C, Klein J, Eyvindson K, Burgas D, Snäll T, Mönkkönen M, Astrup R, Di Fulvio F, Forsell N, Hartikainen M, Uhl E, Poschenrieder W,  Antón-Fernández, C. 2023. What drives forest multifunctionality in central and northern Europe? Exploring the interplay of management, climate, and policies. Ecosystem services 64: e101575.

Burner RC, Stephan JG, Drag L, Potterf M, Birkemoe T, Siitonen J, Müller J, Ovaskainen O, Sverdrup-Thygeson A, Snäll T, 2023. Alternative measures of trait-niche relationships: a test on dispersal traits in saproxylic beetles. Ecology & Evolution 13: e10588.

Blattert C, Mönkkönen M, Burgas D, Di Fulvio F, Toraño Caicoya A, Vergarechea M, Klein J, Hartikainen M, Antón-Fernández C, Astrup R, Emmerich M, Forsell N, Lukkarinen J, Lundström J, Pitzén S, Poschenrieder W, Primmer E, Snäll T, Eyvindson K. 2023.  Climate targets in European timber-producing countries conflict with goals on forest ecosystem services and biodiversity. Communications Earth & Environment 4: Article 119.

Triviño M, Morán-Ordoñez A, Eyvindson K, Blattert C, Burgas D, Repo A, Pohjanmies T, Brotons L, Snäll T, Mönkkönen M. 2023. Future supply of boreal forest ecosystem services is driven by management rather than by climate change. Global Change Biology 29:1484-1500.

Basiri A, Snäll T, Halvorsen T. 2023. Doctoral Thesis Review. Norwegian Journal of Geography 77: 130-132.

Drag L, Burner RC, Stephan JG, Birkemoe T, Dörfler I, Gossner MM, Magdon P, Ovaskainen O, Potterf M, Schall P, Snäll T, Sverdrup-Thygeson A, Weisser W, Müller J. 2023. High-resolution 3D forest structure explains ecomorphological trait variation in assemblages of saproxylic beetles. Functional Ecology  37: 150-161.


Mazziotta A, Lundström J, Forsell N, Moor H, Eggers J, Subramanian N, Aquilué N, Morán-Ordóñez A, Brotons L, Snäll T. 2022. More future synergies and less trade-offs between forest ecosystem services with natural climate solutions instead of bioeconomy solutions. Global Change Biology 28: 6333-6348.

Moor H, Eggers J, Fabritius H, Forsell N, Henckel L, Bradter U, Mazziotta A, Nordén J, Snäll T. 2022. Rebuilding green infrastructure in boreal production forest given future global wood demand. Journal of Applied Ecology 59: 1659-1669.

Eggers J, Lundström J, Snäll T, Öhman K. 2022. Balancing wood production and biodiversity in intensively managed boreal forest. Scandinavian Journal of Forest Research 37: 213-225.

Burner RC, Drag L, Stephan JG, Wetherbee R, Birkemoe T, Müller J, Potterf M, Skarpaas O, Snäll T, Sverdrup-Thygeson A. 2022. Functional structure of European forest beetle communities is enhanced by rare species. Biological Conservation 267: 109491. 

Leitão PJ, Toraño Caicoya A, Dahlkamp A, Guderjan L, Griesser M, Haverkamp P, Nordén J, Snäll T, Schröder B. 2022. Impacts of forest management on forest bird occurrence patterns – a case study in a region in Central Europe. Frontiers in Forests and Global Change 5: 786556.

Blattert C, Eyvindson K, Hartikainen M, Burgas D, Potterf M, Lukkarinen J, Snäll T, Toraño Caicoya A, Mönkkönen M. 2022. Sectoral policies cause incoherence in forest management and ecosystem service provisioning. Forest Policy and Economics 136: 102689.


Snäll T, Triviño M, Mair L, Bengtsson J, Moen J. 2021. High rates of short-term dynamics of forest ecosystem services. Nature Sustainability 4: 951-957.

Löbel S, Schröder B, Snäll T. 2021. Projected shifts in deadwood bryophyte communities under national climate and forestry scenarios benefit large competitors and impairing small species. Journal of Biogeography 48: 3170-3184.

Bradter U, Ozgul A, Griesser M, Layton-Matthews K, Eggers J, Singer A, Sandercock BK, Haverkamp PJ, Snäll T. 2021. Habitat suitability models based on opportunistic citizen science data - evaluating forecasts from alternative methods versus an individual-based model. Diversity and Distributions 27: 2397-2411.

Hahn T, Eggers J, Subramanian N, Toraño Caicoya A, Uhl E, Snäll T. 2021. Specified resilience value of alternative forest management adaptations to storms. Scandinavian Journal of Forest Research 36: 585-597. 

Burner RC, Stephan JG, Drag L, Birkemoe T, Müller J, Snäll T, Ovaskainen O, Potterf M, Siitonen J, Skarpaas O, Doerfler I, Gossner MM, Schall P, Weisser WW, Sverdrup-Thygeson A. 2021. Traits mediate niches and co-occurrences of forest beetles in ways that differ among bioclimatic regions. Journal of Biogeography 48: 3145-3157.

Lauria P, Forsell N, Di Fulvio F, Snäll T, Havlik P. 2021. Material substitution between coniferous, non-coniferous and recycled biomass – Impacts on forest industry raw material use and regional competitiveness. Forest Policy and Economics 132: 102588.

Burner RC, Birkemoe T, Stephan JG, Drag L, Müller J, Ovakainen O, Potterf M, Skarpaas O, Snäll T, Sverdrup-Thygeson A. 2021. Choosy beetles: how host trees and boreal forest naturalness may determine dead wood beetle communities. Forest Ecology and Management 487: 119023.

Moor H, Nordén J, Penttilä J, Siitonen J, Snäll T. 2021. Long-term effects of colonization-extinction dynamics of generalist versus specialist wood-decaying fungi. Journal of Ecology 109: 491-503.


Jonsson M, Bengtsson J, Gamfeldt L, Moen J, Snäll T. 2020. Stand age and climate influence forest ecosystem service delivery and multifunctionality. Environmental Research Letters 15: 0940a8.

Henckel L, Bradter U, Jönsson M, Isaac NJB, Snäll T. 2020. Assessing the usefulness of Citizen Science Data for habitat suitability modelling: opportunistic reporting versus sampling based on a systematic protocol. Diversity and Distributions 26: 1276-1290.

Dawson SK, Berglund H, Ovaskainen O, Snäll T, Jonsson BG, Jönsson M. 2020. Fungal functional traits and red-listed species occurrence of set-aside and natural isolates converge over time, without change in species richness. Biological Conservation 251: 108789.

Eriksson O, Forsell N, Eggers J, Snäll T. 2020. Downscaling of long-term global scenarios to regions with a forest sector model. Forests 11: 500.

Nordén J, Harrison P, Mair L, Siitonen J, Lundström A, Kindvall O, Snäll T. 2020. Occupancy versus colonisation-extinction models for projecting population trends at different spatial scales. Ecology and Evolution 10: 3079-3089.

Nordén J, Abrego N, Boddy L, Bässler C, Dahlberg A, Halme P, Hällfors M, Maurice S, Menkis A, Miettinen O, Makipää R, Ovaskainen O, Penttilä R, Saine S, Snäll T, Junninen K. 2020. Ten principles for conservation translocations of threatened wood-inhabiting fungi. Fungal Ecology 44: 100919.

Eggers J, Räty M, Öhman K, Snäll T. 2020. How well do stakeholder-defined forest management scenarios balance economic and ecological forest values? Forests 11: 86.

Jönsson M, Snäll T. 2020. Ecosystem service multifunctionality of low-productivity forests and implications for conservation and management. Journal of Applied Ecology 57: 695-706.


Jonsson M, Bengtsson J, Gamfeldt L, Moen J, Snäll T. 2019. Levels of forest ecosystem services depend on specific mixtures of commercial tree species. Nature Plants 5: 141-147.

Ranius T, Snäll T, Nordén J. 2019. To understand the effect of the spatial configuration of deadwood habitats requires more focus on occurrence patterns and spatial population dynamics. Conservation Biology 33: 1205-1207.

Fabritius H, Singer A, Pennanen J, Snäll T. 2019. Estimation of metapopulation colonization rates from disturbance history and occurrence pattern data. Ecology 100: e02814.

Singer A, Bradter U, Fabritius H, Snäll T. 2019. Dating past colonization events to project future species distributions. Methods in Ecology and Evolution 10: 471-480.


Mair L, Jönsson M, Räty M, Bärring L, Strandberg G, Lämås T, Snäll T. 2018. Land use changes could modify future negative effects of climate change on old-growth forest indicator species. Diversity and Distributions 24: 1416-1425.

Bradter U, Mair L, Jönsson M, Knape J, Singer A, Snäll T. 2018. Can opportunistically-collected Citizen Science data fill a data gap for habitat suitability models of less common species? Methods in Ecology and Evolution 9: 1667-1678.

Löbel S, Mair L, Lönnell N, Schröder B, Snäll T. 2018. Biological traits explain bryophyte species distributions and responses to forest fragmentation and climatic variation. Journal of Ecology 106: 1700-1713.

Ruete A, Jönsson M, Snäll T. 2018. Conservation benefits of implementing international Aichi protection and restoration targets for future epiphyte metapopulations. Journal of Applied Ecology 55: 118-128.


Ronnås C, Werth S, Ovaskainen O, Várkonyi G, Scheidegger C, Snäll T. 2017. Discovery of long-distance gamete dispersal in a lichen-forming ascomycete. New Phytologist 216: 216-226.

Snäll T, Johansson V, Jönsson M, Ortiz C, Hammar T, Caruso A, Svensson M, Stendahl J. 2017. Transient trade-off between climate benefit and biodiversity of harvesting stump for bioenergy. Global Change Biology Bioenergy 9: 1751-1763.

Pohjanmies T, Triviño M, Le Tortorec E, Mazziotta A, Snäll T, Mönkkönen M. 2017. Impacts of forestry on boreal forests: An ecosystem services perspective. Ambio 46: 743-755.

Mair L, Harrison PJ, Räty M, Bärring L, Strandberg G, Snäll T. 2017. Forest management could counteract distribution retractions forced by climate change. Ecological Applications 27: 1485-1497.

Ruete A, Snäll T, Jonsson BG, Jönsson M. 2017. Long-term effects of transient anthropogenic edges and forest fragment size on the occupancy of deadwood-dwelling fungi. Journal of Applied Ecology 54: 1142-1151.

Jönsson M, Ruete A, Kellner O, Gunnarsson U, Snäll T. 2017. Will forest conservation areas protect functionally important diversity of fungi and lichens over time? Biodiversity and Conservation 26: 2547-2567.

Belinchón R, Harrison PJ, Mair L, Várkonyi G and Snäll T. 2017. Local epiphyte establishment and future metapopulation dynamics in landscapes with different spatio-temporal properties. Ecology 98: 741-750.

Mair L, Harrison PJ, Jönsson M, Löbel L, Nordén J, Siitonen J, Lämås T, Lundström A, Snäll T. 2017. Evaluating citizen science data for forecasting species responses to national forest management. Ecology and Evolution 7: 368-378.

Jonsson M, Snäll T, Asplund J, Clemmensen KE, Dahlberg A, Kumordzi BB, Lindahl BD, Oksanen J, Wardle DA. 2017. Divergent responses of β-diversity among organism groups to a strong environmental gradient. Ecosphere 7: e01535.


Ruete A, Snäll T, Jönsson M. 2016. Dynamic anthropogenic edge effects on the distribution and diversity of fungi in fragmented old-growth forests. Ecological Applications 26: 1475–1485.

Maes J, Liquete C, Teller A, Erhard M, Paracchini ML, Barredo JI, Grizzetti B, Cardoso A, Somma F, Petersen J-E, Meiner A, Royo Gelabert E, Zal N, Kristensen P, Bastrup-Birk A, Biala K, Piroddi C, Egoh B, Degeorges P, Fiorina C, Santos-Martín F, Naruševičius V, Verboven J, Pereira HM, Bengtsson J, Gocheva K, Marta-Pedroso C, Snäll T, Estreguil C, San Miguel J, Pérez-Soba M, Grêt-Regamey A, Lillebø A, Abdul Malak D, Condé S, Moen J, Czúcz B, Drakou EG, Zulian G, Lavalle C. 2016. An indicator framework for assessing ecosystem services in support of the EU Biodiversity Strategy to 2020. Ecosystem services 17: 14-23.

Snäll T, Lehtomäki J, Arponen A, Elith J, Moilanen A. 2016. Green Infrastructure design based on spatial conservation prioritization and modeling of biodiversity features and ecosystem services. Environmental Management 57:251–256.

Svensson, M., Caruso, A., Yahr, R., Ellis, C., Thor, G., Snäll,T. 2016. Combined observational and experimental data provide limited support for facilitation in lichens. Oikos 125: 278–283.


Ruete, A., Fritz, Ö. & Snäll, T. 2014. A model for non-equilibrium metapopulation dynamics utilizing data on species occupancy, patch ages and landscape history. Journal of Ecology 102: 678-689.

Johansson, V., Ranius, T. & Snäll, T. 2014. Development of secondary woodland decreases epiphyte metapopulation sizes in wooded grasslands. Biological Conservation 172: 49-55.

Snäll, T., Forslund, P., Jeppsson, T., Lindhe, A., O’Hara, R.B. 2014. Evaluating temporal variation in Citizen Science Data against temporal variation in the environment. Ecography 37: 293-300.

Maes, J., Teller, A., Erhard, M., Murphy, P.,  Paracchini, M.L., Barredo, J.I., Grizzetti, B., Cardoso, A., Somma, F.,  Petersen, J.-E., Meiner, A., Royo Gelabert, E., Zal, N., Kristensen, P., Bastrup-Birk, A., Biala, K., Romao, C., Piroddi, C., Egoh, B., Fiorina, C., Santos, F., Naruševičius, V., Verboven, J., Pereira, H., Bengtsson, J., Kremena, G., Marta-Pedroso, C., Snäll, T., Estreguil, C., San Miguel, J., Braat, L., Grêt-Regamey, A., Perez-Soba, M., Degeorges, P., Beaufaron, G., Lillebø, A., Abdul Malak, D., Liquete, C., Condé, S., Moen, J., Östergård, H., Czúcz, B., Drakou, E.G., Zulian, G., Lavalle, C. 2014. Mapping and Assessment of Ecosystems and their Services – Indicators for ecosystem assessments under Action 5 of the EU Biodiversity Strategy to 2020. European Commission. ISBN: 978-92-79-36161-6.


Gamfeldt, L., Snäll, T., Bagchi, R., Jonsson, M., Gustafsson, L., Kjellander, P.,  Ruiz-Jaen, M.C., Fröberg, M., Stendahl, J., Philipson, C.D., Mikusiński, G., Andersson, E., Westerlund, B., Andrén, H., Moberg, F., Moen, J., Bengtsson, J. 2013. Higher levels of multiple ecosystem services are found in forests with more tree species. Nature Communications 4: Article number 1340.

Johansson, V., Ranius, T. & Snäll, T. 2013. Epiphyte metapopulation persistence after drastic habitat decline and low tree regeneration: time lags and effects of conservation actions. Journal of Applied Ecology 50: 414-422.

Johansson, V., Snäll, T., & Ranius, T. 2013. Estimates of connectivity reveal non-equilibrium epiphyte occurrence patterns almost 180 years after habitat decline. Oecologia 172: 607-615.

Angelstam, P., Roberge, J.-M., Axelsson, R., Elbakidze, M., Bergman, K.-O., Dahlberg, A., Degerman, E., Eggers, S., Esseen, P.-A., Hjältén, J., Johansson, T., Müller, J., Paltto, H., Snäll, T. , Soloviy, I., Törnblom, J. 2013. Evidence-based knowledge versus negotiated indicators for assessment of ecological sustainability: the Swedish Forest Stewardship Council standard as a case study. Ambio 42: 229-240.


Ruete A, Yang W, Bärring L, Stenseth NC & Snäll T. 2012. Disentangling effects of uncertainties on population projections: climate change impact on an epixylic bryophyte. Proceedings of the Royal Society B: 279: 3098-3105.

Löbel, S., Snäll, T. & Rydin, H. 2012. Epiphytic bryophytes near forest edges and on retention trees: reduced growth and reproduction especially in old-growth-forest indicator species. Journal of Applied Ecology 49: 1334-1343.

Johansson V, Ranius T & Snäll T. 2012. Epiphyte metapopulation dynamics are explained by species traits, connectivity and patch longevity. Ecology 93: 235–241.

Ruete A, Wiklund K & Snäll T. 2012. Hierarchical Bayesian estimation of the population viability of an epixylic moss. Journal of Ecology 100: 499-507. (Journal of Ecology's blog)

Zartman CE, Nascimento HEM, Cangani KG & Alvarenga LDP & Snäll T. 2012. Fine-scale changes in connectivity affect the metapopulation dynamics of a bryophyte confined to ephemeral patches. Journal of Ecology 100: 980-986.

Fedrowitz K, Kuusinen M & Snäll T. 2012. Metapopulation dynamics and future persistence of epiphytic lichens in boreal forests. Journal of Applied Ecology 49: 493–502.

Lankia, H., Wallenius, T., Várkonyi, G., Kouki, J. & Snäll, T. 2012. Forest fire history, aspen, and goat willow in a Fennoscandian old-growth landscape: are current population structures a legacy of historical forest fires? Journal of Vegetation Science 23: 1159-1169.


Snäll T, Kindvall O, Nilsson J, Pärt T. 2011. Evaluating citizen-based presence data for bird monitoring. Biological Conservation 144: 804-810.

Roberge J-M, Johansson S, Wulff S & Snäll T. 2011. Edge creation and tree dieback influence the patch-tracking metapopulation dynamics of a red-listed epiphyte. Journal of Applied Ecology 48: 650-658.

Paltto H, Nordberg A, Nordén B & Snäll T. 2011. Development of secondary woodland in oak wood pastures reduces the richness of red-listed epiphytic lichens. PlosONE 6: e24675.

Björkman C, Johansson H & Snäll T. 2011. Spatial distribution of competing insect predators: possible roles of intraguild predation and the surrounding habitat. Basic and Applied Ecology 12: 516-522.


Caruso, A., Thor, G. & Snäll T. 2010. Colonization-extinction dynamics of epixylic lichens along a decay gradient in a dynamic landscape. Oikos 119: 1947-1953.

Orians C.M., Hochwender C.G., Fritz R.S., and Snäll T. 2010. Growth and chemical defense in willow seedlings: trade-offs are transient. Oecologia 163: 283-290.

Johansson, V., Snäll, T., Johansson, P & Ranius, T. 2010. Detection probability and abundance estimation of epiphytic lichens based on height-limited surveys. Journal of Vegetation Science 21: 332-341.


Snäll, T., Benestad, R.E. & Stenseth, N.C. 2009. Expected future plague levels in a wildlife host under different scenarios of climate change. Global Change Biology, 15, 500-507.

Löbel, S., Snäll, T. & Rydin, H. 2009. Mating system, reproduction mode and diaspore size affect metacommunity dynamics. Journal of Ecology 97, 176-185.

Jonsson P, Sandgren F, Lindberg E & Snäll T. 2009. Towns and rural industrialisation in Sweden 1850-1890: a spatial statistical approach. Scandinavian Economic History Review 57:229-251.


Ben Ari, T., Gershunov, A. Gage, K.L., Snäll, T, Ettestad, P, Kausrud, K.L. & Stenseth, N.C. 2008. Human plague in US: the importance of regional and local climate. Biology Letters, 4, 737-740.

Chi, C.N., Elfström, L., Shi, Y., Snäll, T., Engström, Å. & Jemth, P. 2008. Reassessing a sparse energetic network within a single protein domain. Proceedings of the National Academy of Sciences of the United States of America, 105: 4679-4684.

Snäll, T., O’Hara, R.B., Ray, C. & Collinge, S.K. 2008. Climate-driven spatial dynamics of plague among prairie dog colonies. The American Naturalist, 171: 238-248.


Ringvall, A., Snäll, T., Ekström, M. & Ståhl, G. 2007. Unrestricted guided transect sampling for surveying sparse species. Canadian Journal of Forest Research, 37: 2575-2586.

Snäll, T., O’Hara, R.B. & Arjas, E. 2007. A mathematical and statistical framework for modelling dispersal. Oikos 116:966-974.

Kuparinen, A., Snäll, T., Vänskä, S. O’Hara, R.B. 2007. The role of model selection in describing stochastic ecological processes. Oikos 116:1037-1050.


Löbel, S. Snäll, T., and Rydin, H. 2006. Species richness patterns and metapopulation processes - evidence from epiphyte communities in boreo-nemoral forests. Ecography 29:169-182.

Löbel, S. Snäll, T., and Rydin, H. 2006. Metapopulation processes in epiphytes inferred from patterns of regional distribution and local abundance in fragmented forest landscapes. Journal of Ecology 94:856-868.


Snäll, T., Pennanen, J., Kivistö, L. and Hanski, I. 2005.Modelling epiphyte metapopulation dynamics in a dynamic forest landscape. Oikos 109, 209-222.

Snäll, T., Ehrlén, J. and Rydin, H. 2005.Colonization-extinction dynamics of an epiphyte metapopulation in a dynamic landscape. Ecology 86: 106-115.


Snäll, T., Hagström, A., Rudolphi, J. and Rydin, H. 2004. Distribution pattern of the epiphyte Neckerapennata on three spatial scales - importance of past landscape structure, connectivity and local conditions.Ecography 27: 757-766.

Snäll, T., Fogelqvist, J., Ribeiro Jr., P. J. and Lascoux, M. 2004.Spatial genetic structure in two congeneric epiphytes with different dispersal strategies analysed by three different methods. Molecular Ecology 13: 2109-2119.


Snäll, T., RibeiroJr, P. J. and Rydin, H. 2003. Spatial occurrence and colonisations in patch-tracking metapopulations of epiphytic bryophytes: local conditions versus dispersal. Oikos 103: 566-578.

Snäll, Tord. 2003. Distribution Patterns and Metapopulation Dynamics of Epiphytic Mosses and Lichens. Acta UniversitatisUpsaliensis.Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology, 918.36 pp. Uppsala.


Snäll T. and Jonsson B. G. 2001. Edge effects on six polyporous fungi used as indicators in forest fragments. Ecological Bulletins 49: 55-62.

National publications

Moor H, Siitonen J, Snäll T. 2018. Predicted forest beetle distributions in Dalarna. ArtDatabanken Rapporterar 20. ArtDatabanken, SLU. Uppsala.

Hedenås H, Snäll T, Adler S, Gardfjell H, Lämås T, Mair L. 2017. Miljöövervakning, medborgardata och modellbaserade kartor. I Skog & Mark Tema miljöövervakningen då – nu – sedan. Sid 28-32. Naturvårdsverket. Arkitektkopia AB.

Jönsson M, Bradter U, Mair L, Snäll T. 2017. Medborgarforskning om biologisk mångfald. Biodiverse 22(2): 14-15.

Jönsson M, Snäll T, Mair L. 2016. The role of Swedish LifeWatch in biodiversity citizen science. In Swedish LifeWatch – a national e-infrastructure for biodiversity data. Summary Report 2010-2016. Pp 57-59. ArtDatabanken.

Jönsson M, Ruete A & Snäll T. 2016. Kan frivilliga avsättningar i skogen fungera som ”livbåtar” för rosenticka och lunglav? Biodiverse 21: 22-23.

Eriksson A, Snäll T, Harrison PJ. 2015. Analys av miljöförhållanden – SKA 15. Swedish Forest Agency, Report 11.

Jönsson M, Ruete A, Gunnarsson U, Kellner O, Snäll T. 2015. Övervakning av värdefulla skogsbiotoper – en utvärdering av extensivmetoden efter 10 år. ArtDatabanken Rapporterar 18. ArtDatabanken, SLU, Uppsala.

Johansson V, Caruso A & Snäll T. 2013. Vedlevande lavarters populationsutveckling i fragmenterade landskap: effekten av olika scenarier av stubbskörd. Swedish Energy Agency project Report.

Snäll T, Moen J, Berglund H, Bengtsson J. 2014. Mapping and assessment of ecosystems and their services – the Swedish forest pilot. Rapport 6626, Naturvårdsverket. Bromma.

Johansson V, Ranius T & Snäll T. 2013. Brist på gamla ekar hotar lavar. Svensk Botanisk Tidskrift 6: 344-349.

Jönsson M & Snäll T. 2013. Citizen science i gränslandet mellan vetenskap och folkbildning. Biodiverse 1: 28-29.

Fedrowitz K & Snäll T. 2012. Många stora aspar ger lavarna en chans. Notiser från SLU 2012:June.

Snäll, T. & Roberge, J-M. 2011. Buffertzon kan skydda rödlistade arter. Notiser från SLU, 2.

Snäll, T. & Kellner, O. 2003. Utvärdering av metod för övervakning av skogsbiotoper. Länsstyrelsen Gävleborgs län. Rapport 2003:15.

Naturvårdsverket, by Snäll, T. 1999. In Handbok för miljöövervakning: Extensiv övervakning av skogsbiotopers (-bestånds) innehåll med inriktning mot biologisk mångfald. (

Snäll, T. 1997. Regional övervakning av skogsområden i Dalarna. Miljövårdsenheten, Länsstyrelsen Dalarna


Professor at the Swedish Species Information Centre; SLU Artdatabanken Terrestra ekosystem 1
Telephone: +4618672612, +46767662612